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Strut-Autografting with and without Osteogenic Protein-1 A Preliminary Study of a Canine Femoral Head Defect Model
Michael A. Mont, MD; Lynne C. Jones, PhD; John J. Elias, PhD; Nozomu Inoue, MD, PhD; Taek-Rim Yoon, MD; Edmund Y.S. Chao, PhD; David S. Hungerford, MD
View Disclosures and Other Information
Investigation performed at the Department of Orthopaedic Surgery, The Johns Hopkins University School of Medicine, Baltimore, Maryland
Michael A. Mont, MD
Institute for Advanced Orthopaedics, Sinai Hospital, 2411 West Belvedere Avenue, Suite 102, Baltimore, MD 21215. E-mail address: rhondamont@aol.com

Lynne C. Jones, PhD
John J. Elias, PhD
Nozomu Inoue, MD, PhD
Taek-Rim Yoon, MD
Edmund Y.S. Chao, PhD
David S. Hungerford, MD
Department of Orthopaedic Surgery, The Johns Hopkins University School of Medicine, Good Samaritan Professional Building, 5601 Loch Raven Boulevard, Baltimore, MD 21239

No benefits in any form have been received or will be received from a commercial party related directly or indirectly to the subject of this article. Funds were received in total or partial support of the research or clinical study presented in this article. The funding source was Grant 95-018 from the Orthopaedic Resident and Education Foundation. Stryker Biotech (Hopkinton, Massachusetts) supplied the osteogenic protein-1.

The Journal of Bone & Joint Surgery.  2001; 83:1013-1022 
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Abstract

Background: Osteonecrosis of the femoral head frequently leads to collapse of the articular cartilage and to disabling osteoarthritis, which ultimately may necessitate joint arthroplasty. One treatment method that has had moderate success is the so-called trapdoor approach, which involves excavation of diseased (necrotic) bone followed by bone-grafting. Augmentation of this procedure with various growth and differentiation factors may improve the outcome. We developed a canine model that mimics the clinical situation with trapdoor bone-grafting. The objective of this study was to evaluate the effect of the addition of osteogenic protein-1 on healing following the trapdoor procedure with strut-autografting.

Methods: Thirty-four skeletally mature dogs were used in the experiment. After capsulotomy, a trapdoor was created in the anterolateral surface of the femoral head and a 2-cm-diameter subchondral area of bone was removed. In the phase-I experiments, seven dogs had no treatment of the defect (Group I) and nine dogs were treated with strut-grafting (Group II). In phase II, the procedure was modified by collapsing the trapdoor into the created defect intraoperatively in eighteen dogs, which were divided into three equal groups: six untreated defects were left collapsed (Group III), six were treated with bone graft (Group IV), and six were treated with bone graft augmented with osteogenic protein-1 (Group V).

Results: Three of the seven femoral heads in Group I (untreated defect) and one of the nine heads in Group II (grafting without collapsing of the trapdoor) had evidence of cartilage collapse. Inspection of sagittal slices and radiographs revealed an unfilled residual defect in all Group-I heads, whereas all Group-II heads were well healed. The mean normalized stiffness value was significantly larger in Group II than it was in Group I. On visual inspection, depression was noted in all of the femoral heads in Group III (untreated defect; trapdoor left collapsed). In both Group IV and Group V (grafting without and with osteogenic protein-1), the trapdoor cartilage appeared to be essentially normal. Groups IV and V had more radiographic healing than did Group III. The defects in Group V (grafting with osteogenic protein-1) healed faster radiographically than did those in Group IV (grafting without osteogenic protein-1).

Conclusions: Moderate-to-excellent healing was seen both radiographically and biomechanically by four months in the groups treated with grafting, with and without osteogenic protein-1, whereas untreated defects did not heal.

Clinical Relevance: Symptomatic osteonecrosis of the femoral head is a clinical challenge. The animal model in the current study is a useful tool for the evaluation of methods to treat osteonecrosis of the femoral head. Studies investigating additional time-periods between implantation of osteogenic protein-1 and assessment of results as well as different doses of osteogenic protein-1 are warranted.

Figures in this Article
    Osteonecrosis of the femoral head leads to loss of mechanical stability, collapse of the articular surface, and degenerative joint disease. The majority of patients with osteonecrosis are first seen when the disease is in an advanced stage—that is, after collapse of the femoral head, a condition that may necessitate total hip arthroplasty. Moreover, reports describing the long-term results of hip replacements have commonly described a high frequency of failure in patients with osteonecrosis1-6, underscoring the importance of developing successful joint-preserving procedures for patients with this disease.
    If a diagnosis is made prior to collapse of the femoral head, treatment alternatives include core decompression, osteotomy, and various vascularized and nonvascularized bone-grafting methods7-19. Most reports in the literature do not present conclusive data to support the effectiveness of any one surgical procedure in preventing segmental collapse of the femoral head.
    Bone-grafting is performed with the aims of removing diseased bone, speeding up the revascularization process, and providing structural reinforcement during healing in the hope of preserving the contour of the femoral head4,7-9,11,12,14-16,18,20. Bone-grafting may be useful in the treatment of early osteonecrosis (stages I and II, according to the system of Ficat and Arlet21, whereas it is less effective in the later, symptomatic stages (Ficat and Arlet stages III and IV). Some authors have initiated the clinical use of cortical strut and cancellous autografting through a so-called trapdoor in the femoral head for the treatment of late stages of osteonecrosis (Ficat and Arlet stages IIC, III, and early IV) when the only other surgical alternative would be total hip arthroplasty8,9,11,12,19. Although the five-year results in one study were good or excellent in twenty of twenty-four stage-III hips, enhancement of the healing process could make this procedure even more efficacious12.
    Basic and clinical research has demonstrated the efficacy of various growth and differentiation factors (bone morphogenetic proteins, interleukins, and angiogenic growth factors) in healing bone defects13,20,22-39, as an alternative or adjunct to bone grafts. Recently, studies have been performed to assess the role of growth and differentiation factors in the treatment of osteonecrosis13,29,33.
    Rigorous testing is needed in order to evaluate the different graft-treatment regimens available for patients with osteonecrosis. Unfortunately, there is no entirely satisfactory animal model for atraumatic human osteonecrosis33,40-45. In the present study, we used a femoral head defect model that directly mimics the excavated bone in the femoral head in patients with osteonecrosis treated by the trapdoor approach. The treatment of the animals was similar to that of patients, with the dead bone completely removed by curettage. We present the results of autogenous bone-grafting with and without the use of a bone morphogenetic protein (osteogenic protein-1) in this model.
     
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    +Fig. 1:Diagram illustrating the trapdoor technique. The trapdoor is made in the femoral head cartilage (a), after which a 2-cm-diameter area of the femoral head bone is excavated (b). In Group I, the defect was left unfilled (c) and the trapdoor cartilage segment was replaced (d). In Group III, this cartilage segment was collapsed. In Groups II, IV, and V, iliac crest bone graft was harvested (e) and utilized as cancellous bone chips and cortical bone struts (f, g, and h), and the trapdoor was replaced (i). In Group V, osteogenic protein-1 was added.
     
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    +Fig. 2-A:Each femur was secured within a fixation jig fixed to the mechanical testing machine. The fixation jig allowed two degrees of translational adjustment and three degrees of rotational adjustment at the base of the embedded femur. Each test site was aligned with the indentor fixed to the loading actuator.
     
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    +Fig. 2-B:A standard technique was developed to identify the test positions. A circle was mapped onto the femoral head with one point tangential to the plane (plane A) through the posterior edge of the greater trochanter (arrow) and the fovea and a second point tangential to the plane (plane B) initiating at the fovea and perpendicular to the first plane. The test sites were distributed around the perimeter of the circle. For Groups I and II, the posterior-medial (2), anterior-medial (3), anterior-lateral (5), posterior-lateral (6), and center (7) test sites were used. For Groups III, IV, and V, the center site was used again, along with a posterior site (1) and an anterior site (4).
     
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    +Fig. 3-A:Figs. 3-A through 3-E Representative radiographs from the five study groups. Fig. 3-A Group I demonstrated no healing.
     
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    +Fig. 3-B:Group II demonstrated excellent healing.
     
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    +Fig. 3-C:Minimal radiographic healing with collapse of the femoral head was seen in Group III.
     
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    +Fig. 3-D:Moderate radiographic healing without collapse of the femoral head was seen in Group IV.
     
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    +Fig. 3-E:Excellent radiographic healing without collapse of the femoral head was seen in Group V.
     
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    +Fig. 4-A:A histological section through the cartilage of the collapse model (Group III) illustrates the irregularity of the cartilage surface (Goldner trichrome, 25).
     
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    +Fig. 4-B:A histological section of a femoral head treated with bone-grafting (Group IV). There was persistence of the defect where the trapdoor had been made. Fibrous tissue is evident between the ends of the cartilage, where bone has grown underneath the cartilage (Goldner trichrome, 25).
     
    Anchor for JumpAnchor for JumpTABLE I:  Normalized Stiffness Values*
    *The values are given as the mean and the standard deviation.
    Test SiteGroup IGroup IIGroup IIIGroup IVGroup V
    Anterior-lateral 0.8±0.31.2±0.6
    Anterior-medial0.9±0.31.1±0.6
    Center1.1±0.41.3±0.50.9±0.31.0±0.20.9±0.2
    Posterior-lateral0.9±0.31.0±0.4
    Posterior-medial0.8±0.21.0±0.3
    Anterior1.0±0.20.9±0.21.0±0.4
    Posterior0.9±0.21.1±0.20.8±0.4
    Thirty-four skeletally mature, heartworm-free, mongrel dogs weighing 28.9 to 40.9 kg were used. There were twenty-three female and eleven male animals. The animal protocol was reviewed and approved by the institutional animal care and use committee. A pelvic radiograph was made of each dog to screen for signs of bone disease or hip dysplasia and to confirm skeletal maturity.

    Experimental Design (Fig. 1)

    In phase I, sixteen dogs were used to create the defect model. Seven defects were left untreated (Group I), and nine were treated with the strut-grafting technique (Group II). In phase II, the surgical procedure was modified, in order to provide a more reproducible collapse of the articular surface, by allowing the articular cartilage of the trapdoor to collapse into the untreated defect intraoperatively in eighteen dogs. The trapdoor was left collapsed in six femoral heads (Group III), the collapsed segment was elevated with bone-grafting in six (Group IV), and the collapsed segment was elevated with a combination of bone-grafting and osteogenic protein-1 in six (Group V). All animals in all five groups were killed at four months postoperatively.

    Surgical Procedures

    Dogs were tranquilized with acetylpromazine maleate (1 mg/kg intramuscularly) followed by anesthesia with Ethrane (enflurane), nitrous oxide, and oxygen. With use of aseptic technique, the right or left hip (chosen randomly) of each experimental animal was exposed through an anterolateral approach46. In this approach, the major vessels supplying the head (the lateral circumflex branch of the profunda femoris) are preserved. The hip capsule was exposed, and a capsulotomy was performed without dislocating the hip to expose the femoral head and neck. A 1-cm-diameter trapdoor composed of articular cartilage and its underlying subchondral bone was created on the anterolateral aspect of the femoral head (Fig. 1). The trapdoor was removed, and a mass of trabecular bone was excised from the femoral head to create a spherical defect 2 cm in diameter. The size of the trapdoor corresponds to the size (70% to 80% of the femoral neck diameter) that has been utilized in trapdoor procedures in human clinical studies9,12,14,19. Bone was removed until the subchondral bone and cartilage adjacent to the defect was 2 mm thick. After the grafting procedures, the cartilage flap was replaced flush with the surrounding articular surface and held in place by two 6-0 sutures and by closing the capsule with two 6-0 sutures. The hip abductors were then repaired.
    In phase II, the surgical procedure described above was modified after the bone was removed. All animals had intraoperative collapse of the trapdoor (consisting of articular cartilage and its underlying subchondral bone) by application of light pressure to the trapdoor with use of an 8-mm-diameter bone tamp until the trapdoor was depressed into the defect 3 mm. In Group III, this segment was left collapsed. In Groups IV and V, the trapdoor was elevated, the defect was filled with bone graft without or with osteogenic protein-1, and the trapdoor was replaced, flush with the surrounding cartilage.
    After all procedures, the animals were permitted to recover from the general anesthesia and were allowed free activity. The animals were observed daily for wound-healing and gait assessment. All animals received one preoperative dose and two postoperative doses of antibiotics (gentamicin, 4 mg/kg). When grafting was to be performed, autogenous corticocancellous bone was taken from the ilium through the same incision and was utilized to pack the defect. For animals to be treated with osteogenic protein-1 (Stryker Biotech, Hopkinton, Massachusetts), an equal volume of this material was mixed with the autogenous cancellous bone graft (250 mg osteogenic protein-1/g of bone) and packed into the defect after the cortical struts were placed.

    Load-Bearing

    The ground-reaction forces were determined before and at eight and sixteen weeks after the surgery to assess functional walking in the phase-I studies. Dogs were led along a 7-m-long and 1-m-wide platform. Velocity was maintained consistently and was measured as the time between forelimb and hindlimb strikes. A force-plate (model OR-6-6; Advanced Mechanical Technology, Newton, Massachusetts) was mounted level with the top of the platform. As the dog struck the force-plate, an analog signal was digitized and processed by specialized software designed to interpret canine gait47,48. Each individual force-plate-analysis session consisted of a minimum of six trials for both the left and the right hindlimb. Vertical ground-reaction force was determined for each trial, and the mean value was calculated. Data were normalized to body weight. Differences between the mean values in Groups I and II and time-sequential changes in mean values in each group were analyzed.

    Radiographic Analysis

    Anteroposterior and lateral radiographs were made preoperatively, immediately postoperatively, and at one, two, and three months postoperatively. After the animals were killed (four months postoperatively), final radiographs of the dissected femoral heads were made with a Faxitron machine (Hewlett-Packard, Palo Alto, California).
    The radiographs were evaluated by four clinicians who were blinded to the animal groups and who used a scoring system in which filling of the defect was graded as none or scant, minimal, moderate, or excellent. None or scant healing of the defect denoted minimal bone formation (filling <25% of the defect) composed mostly of noncontiguous areas of minimal density. Minimal healing indicated mostly contiguous areas of normal density that filled 25% to 75% of the defect. Moderate healing encompassed normal density in 76% to 95% of the defect. Excellent healing denoted normal density in >95% of the defect.

    Harvesting of Femoral Heads

    Specimens from the wound were taken for aerobic and anaerobic bacterial cultures after the animals were killed with an overdose of sodium pentobarbital (200 mg/kg). Femoral samples were then harvested and were examined macroscopically and photographed. The bone was then dissected free and processed for mechanical testing, followed by histological examination.

    Biomechanical Testing (Figs. 2-A and 2-B)

    After the animals were killed, both femora were excised from each dog and were placed in a bath of ice and lactated Ringer solution until testing. The femora were cut approximately 5 cm distal to the neck and were embedded in a cube of low-melting-point metal (Cerro-Bend Metals, Belfonte, Pennsylvania) at an angle of 30° to the vertical, to position the weight-bearing surface at approximately the highest point of the embedded femur. Indentation tests were performed after removal of the articular cartilage from the specific test sites with a scalpel to expose the subchondral bone. A line was drawn along the femoral head between the osseous prominence on the posterior edge of the greater trochanter and the fovea. A second line originating at the fovea was drawn perpendicular to the first line. A circular template with a 0.5-in (1.27-cm) diameter was aligned with the lines. The center of the circle marked the center test site. For the Group-I and Group-II specimens, four additional test sites evenly spaced along the circumference of the circle were identified on each femoral head; they were identified as the anterior-medial, anterior-lateral, posterior-medial, and posterior-lateral test sites. This procedure resulted in a testing region that approximately spanned the weight-bearing region of the femoral head. For the specimens in Groups III, IV, and V, only three sites were used, to reduce the variability in the stiffness measurements49-51. The same central test site was used along with one site approximately 0.25 in (0.64 cm) anterior to it and another approximately 0.25 in posterior to it. Each embedded femur was secured within a fixation jig fixed to the loading frame of a mechanical testing machine (Bionix 858; MTS, Eden Prairie, Minnesota) on a base-plate that allowed translation in two dimensions and rotational adjustment in three. The base-plate is a rotational joint within the fixation jig, which was used to position each test site approximately perpendicular to the coding actuator. A compressive load was applied to each test site at a rate of 2 mm/min to a maximum load of 125 N with use of a 3-mm-radius indentor52.
    Preliminary experiments showed a failure strength of nearly 400 N for hollowed-out femoral heads. Each site was tested four times in succession. The stiffness consistently increased from the first through the third test. The maximum stiffness from the third and fourth tests was used for statistical analysis. At each test site, the stiffness value of the surgically altered head was normalized by the stiffness of the untreated, contralateral control. During testing, each femoral head was kept moist with saline solution.

    Histological Study

    After biomechanical testing, the proximal part of the femur was isolated and then sectioned longitudinally and transversely. The undecalcified bone was sequentially dehydrated in increasing concentrations of ethanol. The specimens were then embedded in methylmethacrylate according to the technique of Emmanual et al.53. Each block was sectioned at 200 mm with use of the EXAKT processing system. The specimens were then ground to 70 to 100-mm sections. Next, each block was cut into 5-mm sections with use of an Exakt-System microtome (Exakt Apparatebau, Munich, Germany). The slides were then stained with hematoxylin and eosin), MIBS (Villanueva mineralized bone stain), Goldner trichrome, toluidine blue, and safranin O.

    Data Analysis

    Parametric comparisons of each treated head with the contralateral, intact head were made with use of the Student t test for matched pairs. Comparisons between independent groups (untreated defects, defects treated with grafting, and defects treated with grafting and osteogenic protein-1) were carried out with use of one-way analysis of variance (randomized) with the post hoc Newman-Keuls between-group comparisons test. Nonparametric testing between two independent groups was done with the Kruskal-Wallis test. Comparison between groups with regard to success or failure of healing was performed with the Fisher exact test. For the biomechanical studies, a nested analysis of variance was used to compare the normalized stiffness values between the group treated with grafting and the untreated group. The nested analysis of variance treated each normalized stiffness value as an equivalent data point, regardless of its magnitude. A separate analysis of variance, combined with a Tukey-Kramer test for multiple comparisons, was performed to examine the variations in the stiffness magnitudes among the five test sites in the control femora.
    All animals tolerated the operation well. All wounds healed with no dehiscence, and no other complication, such as hip dislocation or deep infection, was encountered.

    Load-Bearing

    All animals were able to stand unassisted within the first twenty-four-hour period. Ground-reaction forces were determined with a force-plate for the Group-I animals (untreated defects) and the Group-II animals (defects treated with grafting) at eight and sixteen weeks postoperatively. Load-bearing on the treated limb was significantly higher in Group II than it was in Group I (p < 0.01). In Group I, dynamic loading of the involved limb had significantly decreased to a mean of 87% of the preoperative value at eight weeks after the surgery (p < 0.05). The level increased slightly, to a mean of 91%, by sixteen weeks, but the increase was not significant. In Group II, no significant decrease in dynamic loading of the involved limb, as compared with that of the contralateral limb, could be detected at eight or sixteen weeks after the surgery.

    Gross Appearance of the Femoral Heads

    Three of the seven femoral heads in Group I and one of the nine heads in Group II appeared to have irregularities in the articular cartilage and evidence of collapse at the defect site. Macroscopically, the articular cartilage of these four collapsed heads was depressed between 1 and 4 mm. The articular cartilage of the trapdoor and the cartilage adjacent to the trapdoor were depressed, soft, friable, and fissured. All of the other femoral heads in Groups I and II had minimal cartilage irregularities. The perimeter of the trapdoor had a grayish-white appearance, was soft, and appeared to be composed of fibrous tissue. This approximately 0.5-mm band of tissue surrounded essentially normal-appearing, nondepressed cartilage of the trapdoor. The anatomy was restored and was not grossly different between the two groups.
    All of the Group-III animals (collapse of the trapdoor without grafting) had collapse of the femoral head ranging from 1 to 4 mm. The articular cartilage was soft, friable, and yellow and had a rough surface with clefts and fissures.
    In Group IV (collapse of the trapdoor followed by grafting without osteogenic protein-1) and Group V (collapse of the trapdoor followed by grafting with osteogenic protein-1), the trapdoor cartilage as well as the adjacent cartilage overlying the filled defect had an almost normal appearance. There were only minor fibrillations, which for the most part resembled normal hyaline cartilage, in some specimens. The perimeter of the trapdoor in these specimens was similar to that in Groups I and II.

    Radiographic Analysis (Figs. 3-A, 3-B, 3-C, 3-D, and 3-E)

    Postoperatively, all of the Group-I defects were clearly visible and sharply demarcated on standard anteroposterior and lateral radiographs (Fig. 3-A). All of the Group-II femoral heads exhibited a greater healing response (sclerosis across the defect increasing at each time-interval, with no femoral head appearing collapsed; Fig. 3-B) than the Group-I femoral heads (persistence of the lucent defect with minimal sclerotic bridging). By four months, seven of the nine Group-II femoral heads showed excellent radiographic healing of the defect and two showed moderate healing. There was minimal healing of the defect in all seven Group-I animals.
    The radiographs made when the animals were killed revealed minimal healing in all six Group-III femoral heads (Fig. 3-C). The defects in Group V healed faster radiographically than did those in Group IV. Of the six femoral heads in Group IV, four showed excellent healing and two showed moderate healing (Fig. 3-D). In Group V, all six femoral heads demonstrated excellent healing (Fig. 3-E).

    Histological Analysis (Figa. 4-A and 4-B)

    Histological analysis demonstrated fibrillation of the cartilage in all of the femoral heads. In Group III, the trapdoor remained collapsed in all specimens (Fig. 4-A), whereas there were no areas of cartilage depression in Group IV (Fig. 4-B) or V. There was no significant difference in cartilage thickness among the three groups, although Group IV had a slightly greater cartilage width (mean, 212.5 mm) than did Groups V and III (mean, 167 and 166 mm, respectively). These values were not significantly different from the control widths (mean, 185 mm).

    Biomechanical Indentation Testing

    The mean normalized stiffness value was significantly greater (p < 0.05) in Group II than it was in Group I at three of the five indentation test sites (anterior-lateral, anterior-medial, and posterior-medial) (Table I). The nested analysis of variance did not indicate that the test site significantly influenced the normalized stiffness. A separate analysis of variance combined with a Tukey-Kramer analysis for post hoc testing was used to compare the stiffness magnitudes among the test sites of the untreated, control femoral head from each dog. The stiffness values ranged from 397 151 N/mm for the anterior-lateral test site to 1021 228 N/mm for the posterior-medial test site. The two anterior sites were significantly less stiff than the central site and the posterior-medial site. The anterior-lateral site was also significantly less stiff than the posterior-lateral site.
    In contrast to the comparisons between Groups I and II, comparisons among Groups III, IV, and V with use of nested analysis of variance and individual comparisons of the normalized stiffness values at each of the three test sites did not indicate a significant difference in the normalized stiffness values (p = 0.40) (Table I). At each site, the standard deviations of the mean normalized stiffness values were similar to the standard deviations in Groups I and II. The mean stiffness values for the untreated controls in Groups III, IV, and V were 488 108 N/mm at the anterior site, 856 200 N/mm at the center site, and 1076 270 N/mm at the posterior site.

    Femoral Head Bone-Grafting and Enhancement

    To our knowledge, Ganz and Büchler8 were the first to mention cancellous grafting through a window in the femoral neck combined with osteotomies; however, they did not report their results. Various investigators in Japan modified this procedure by using strut grafts through a window in the femoral neck. Rosenwasser et al.14 utilized a similar technique, in which they combined complete evacuation of the femoral head with replacement with cancellous iliac-crest bone; they reported thirteen excellent results in fifteen patients at a mean of twelve years after this procedure. Nonvascularized bone-grafting through a trapdoor in the cartilage of the femoral head was apparently first described by Meyers et al.11. We12 utilized a modification of this procedure and reported twenty good and excellent clinical results at a mean of 4.6 years in twenty-four hips with stage-III disease. Growth and differentiation factors can be added to autogenous bone graft without modifying the procedure substantially. Any enhancement of healing that results may shorten the period of restricted weight-bearing, ensure better compliance with rehabilitation protocols, and hopefully lead to more successful outcomes. These goals led us to perform the present study.

    Osteonecrosis Models

    Over the last seventy years, a great deal of work has been devoted to developing a relevant model for osteonecrosis. These efforts have met with little success. No animal model has produced histological or morphological changes representative of the pathological findings in humans33,40-45. The defect model in the present study is useful as it simulates the defect created in the treatment of late-stage osteonecrosis with the trapdoor approach. The trapdoor procedure, or modifications of it employing various bone-grafting procedures, have been utilized by multiple authors7,9,11,12,14,19. A main feature of the trapdoor procedure in human patients is the removal of all necrotic bone. The evacuated femoral head is left with viable and vascular bone, as it was in our animal model. Knowledge gained from assessment of the healing of this defect after treatment with strut bone-grafting with or without bone morphogenetic protein in dogs could be clinically useful when these same defects are created in patients. In summary, this is an important model for the study of osteonecrosis because it (1) uses the femoral head, (2) involves a subchondral defect, (3) has structural compromise, and (4) is similar to defects made in treatment involving vascularized bone-grafting.

    Present and Related Studies

    In the present study, although all of the Group-I (untreated) defects persisted as seen radiographically, they did not consistently result in collapse of the femoral head; in fact, only three of the seven heads collapsed. As a result of this finding, it was decided that all subsequent femoral heads would be collapsed after the defect had been made at the primary operation and then either left collapsed (Group III) or treated with bone-grafting (Groups IV and V). Both Group IV and Group V (grafting without and with osteogenic protein-1) exhibited excellent healing by four months, as demonstrated by the radiographic and biomechanical studies.
    Mazières29 carried out preliminary experiments with use of recombinant human bone morphogenetic protein-2 in a pig model of osteonecrosis. Three animals underwent bilateral core decompression of the femoral head followed by introduction of a mixture of recombinant human bone morphogenetic protein-2 and blood clot (Genetics Institute, Cambridge, Massachusetts) into the core channel on one side. Three months after the surgery, standard radiographs and magnetic resonance images as well as histological examination revealed evidence of excellent healing of the treated channels compared with that of the open channels. The one-month histological findings in the hips treated with bone morphogenetic protein-2 were comparable with the three-month findings in the hips treated without bone morphogenetic protein-2. These findings are consistent with those of the present study, in which the use of adjunctive bone morphogenetic protein was found to be associated with an acceleration of healing at early time-points.
    Scully et al.33 studied the effects of recombinant human bone morphogenetic protein-2 and vascularized fibular grafting in a canine model of osteonecrosis that had been induced by a combination of soft-tissue dissection and freezing. There were no radiographic differences between the animals treated with bone morphogenetic protein-2 and control animals treated with vascularized fibular graft alone, but quantitative histomorphometry did reveal increased amounts of viable bone at both eight and twelve weeks in the former group when compared with the latter. Scully et al. suggested that revascularization after fibular grafting is accelerated by the addition of bone morphogenetic protein-2. Our study also indicates that growth and differentiation factors enhance the early healing response in an animal model of osteonecrosis.
    In summary, we used an experimental model that is potentially valuable for evaluation of the efficacy of procedures performed to elicit the healing of osseous defects in the femoral head and to prevent collapse of the overlying articular cartilage. Radiographic and biomechanical studies indicated that bone-grafting either with or without osteogenic protein-1 resulted in excellent healing of 2-cm-diameter spherical defects and prevented the displacement and collapse of the trapdoor and the surrounding cartilage. Untreated defects did not heal. Additional studies investigating additional time-periods between implantation of osteogenic protein-1 and assessment of results as well as doses of osteogenic protein-1 are warranted.
    Note: The authors thank David Ruegger and Jaime Kemler for their assistance with the histological preparation. They also acknowledge the following individuals for their technical and support contributions to this investigation: Dawn M. LaPorte, MD, Henri Pierre-Jacques, MD, Ivan H. Pacheco, MD, Rad K. Payman, MD, A.H. Reddi, PhD, Melissa Schlenker, BA, Anthony Valdevitt, PhD, and James F. Wenz, MD.
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    Ganz R, Büchler U. Overview of attempts to revitalize the dead head in aseptic necrosis of the femoral head—osteotomy and revascularization. In: Hungerford DS, editor. The hip: proceedings of the eleventh open scientific meeting of the Hip Society. St. Louis: CV Mosby; 1983. p 296-305 
     
    Itoman M, Yamamoto M: Pathogenesis and treatment of idiopathic aseptic necrosis of the femoral head. Clin Immunol,1989.21: 713-25, 21713  1989 
     
    Mankin HJ: Nontraumatic necrosis of bone (osteonecrosis). N Engl J Med,1992.326: 1473-9, 3261473  1992  [PubMed]
     
    Meyers MH, Jones RE, Bucholz RW, Wenger DR: Fresh autogenous grafts and osteochondral allografts for the treatment of segmental collapse in osteonecrosis of the hip. Clin Orthop,1983.174: 107-12, 174107  1983  [PubMed]
     
    Mont MA, Einhorn TA, Sponseller PD, Hungerford DS: The trapdoor procedure using autogenous cortical and cancellous bone grafts for osteonecrosis of the femoral head. J Bone Joint Surg Br,1998.80: 56-62, 8056  1998  [PubMed]
     
    Mont MA, Jones LC, Einhorn TA, Hungerford DS, Reddi AH: Osteonecrosis of the femoral head. Potential treatment with growth and differentiation factors. Clin Orthop,1998.355(Suppl): 314-35, 355(Suppl)314  1998 
     
    Rosenwasser MP, Garino JP, Kiernan HA, Michelsen CB: Long term followup of thorough debridement and cancellous bone grafting of the femoral head for avascular necrosis. Clin Orthop,1994.306: 17-27, 30617  1994  [PubMed]
     
    Smith KR, Bonfiglio M, Montgomery WJ: Non-traumatic necrosis of the femoral head treated with tibial bone-grafting. A follow-up note. J Bone Joint Surg Am,1980.62: 845-7, 62845  1980  [PubMed]
     
    Steinberg ME, Brighton CT, Corces A, Hayken GD, Steinberg DR, Strafford B, Tooze SE, Fallon M: Osteonecrosis of the femoral head. Results of core decompression and grafting with and without electrical stimulation. Clin Orthop,1989.249: 199-208, 249199  1989  [PubMed]
     
    Sugioka Y, Hotokebuchi T, Tsutsui H: Transtrochanteric anterior rotational osteotomy for idiopathic and steroid-induced necrosis of the femoral head. Indications and long-term results. Clin Orthop,1992.277: 111-20, 277111  1992  [PubMed]
     
    Urbaniak JR, Coogan PG, Gunneson EB, Nunley JA: Treatment of osteonecrosis of the femoral head with free vascularized fibular grafting. A long-term follow-up study of one hundred and three hips. J Bone Joint Surg Am,1995.77: 681-94, 77681  1995  [PubMed]
     
    Yamamoto M, Itoman M, Sagamoto N, Morita M: Strut bone graft for aseptic necrosis of the femoral head: theory and surgical technique. Orthop Surg,1983.34: 902-8, Japanese34902  1983 
     
    Wang EA, Rosen V, D’Alessandro JS, Bauduy M, Cordes P, Harada T, Israel DI, Hewick RM, Kerns KM, LaPan P, Luxenberg DP, McQuaid D, Moutsatsos IK, Nove J, Wozney JM: Recombinant human bone morphogenetic protein induces bone formation. Proc Natl Acad Sci USA,1990.87: 2220-4, 872220  1990  [PubMed]
     
    Ficat RP, Arlet J. Functional investigation of bone under normal conditions. In: Hungerford DS, editor. Ischemia and necrosis of bone. Baltimore: Williams and Wilkins; 1980. p 29-52 
     
    Cook SD, Baffes GC, Wolfe MW, Sampath TK, Rueger DC: Recombinant human bone morphogenetic protein-7 induces healing in a canine long-bone segmental defect model. Clin Orthop,1994.301: 302-12, 301302  1994  [PubMed]
     
    Cook SD, Dalton JE, Tan EH, Whitecloud TS 3rd, Rueger DC: In vivo evaluation of recombinant human osteogenic protein (rhOP-1) implants as bone graft substitute for spinal fusions. Spine,1994.19: 1655-63, 191655  1994  [PubMed]
     
    Cook SD, Salkeld SL, Brinker MR, Wolfe MW, Rueger DC: Use of an osteoinductive biomaterial (rhOP-1) in healing large segmental bone defects. J Orthop Trauma,1998.12: 407-12, 12407  1998  [PubMed]
     
    Gerhart TN, Kirker-Head CA, Kriz MJ, Holtrop ME, Hennig GE, Hipp J, Schelling SH, Wang E: Healing segmental femoral defects in sheep using recombinant human bone morphogenetic protein. Clin Orthop,1993.293: 317-26, 293317  1993  [PubMed]
     
    Johnson EE, Urist MR, Finerman GA: Resistant nonunions and partial or complete segmental defects of long bones. Treatment with implants of a composite of human bone morphogenetic protein (BMP) and autolyzed, antigen-extracted, allogeneic (AAA) bone. Clin Orthop,1992.277: 229-37, 277229  1992  [PubMed]
     
    Khouri RK, Koudsi B, Reddi H: Tissue transformation into bone in vivo. A potential practical application. JAMA,1991.266: 1953-5, 2661953  1991  [PubMed]
     
    Lindholm TC, Lindholm TS, Marttinen A, Urist MR: Bovine bone morphogenetic protein (bBMP/NCP)-induced repair of skull trephine defects in pigs. Clin Orthop,1994.301: 263-70, 301263  1994  [PubMed]
     
    Mazières B: Bone morphogenetic protein and bone necrosis: a perspective. ARCO News,1994.6: 3-5, 63  1994 
     
    Mundy GR: Regulation of bone formation by bone morphogenetic proteins and other growth factors. Clin Orthop,1996.323: 24-8, 32324  1996 
     
    Reddi AH: Symbiosis of biotechnology and biomaterials: applications in tissue engineering of bone and cartilage. J Cell Biochem,1994.56: 192-5, 56192  1994  [PubMed]
     
    Riley EH, Lane JM, Urist MR, Lyons KM, Lieberman JR: Bone morphogenetic protein-2: biology and applications. Clin Orthop,1996.324: 39-46, 32439  1996  [PubMed]
     
    Scully SP, Rizk WS, Seaber AV, Urbaniak JR. Augmentation of subchondral bone formation in AVN with rhBMP-2. Poster presentation at the Annual Meeting of the Orthopaedic Research Society. 1995 Feb 13-16. Orlando, FL 
     
    Toriumi DM, Kotler HS, Luxenberg DP, Holtrop ME, Wang EA: Mandibular reconstruction with a recombinant bone-inducing-factor. Functional, histologic, and biomechanical evaluation. Arch Otolaryngol Head Neck Surg,1991.117: 1101-12, 1171101  1991  [PubMed]
     
    Trippel SB, Coutts RD, Einhorn TA, Mundy GR, Rosenfeld RG: Growth factors as therapeutic agents. J Bone Joint Surg Am,1996.78: 1272-86, 781272  1996 
     
    Urist MR: Bone: formation by autoinduction. Science,1965.150: 893-9, 150893  1965  [PubMed]
     
    Urist MR, Dawson E: Intertransverse process fusion with the aid of chemosterilized autolyzed antigen-extracted allogeneic (AAA) bone. Clin Orthop,1981.154: 97-113, 15497  1981  [PubMed]
     
    Wozney JM: Bone morphogenetic proteins. Prog Growth Factor Res,1989.1: 267-80, 1267  1989  [PubMed]
     
    Yasko AW, Lane JM, Fellinger EJ, Rosen V, Wozney JM, Wang EA: The healing of segmental bone defects, induced by recombinant human bone morphogenetic protein (rhBMP-2). A radiographic, histological, and biomechanical study in rats. J Bone Joint Surg Am,1992.74: 659-70, 74659  1992  [PubMed]
     
    Adams RJ. Experimental surgery and physiology: induced animal models of human disease. In: Sindle MM, Adams RJ, editors. Musculoskeletal system. Baltimore: Williams and Wilkins; 1988. p 41-55 
     
    Gold EW, Fox OD, Weissfeld S, Curtiss PH: Corticosteroid-induced avascular necrosis: an experimental study in rabbits. Clin Orthop,1978.135: 272-80, 135272  1978  [PubMed]
     
    Jones LC, Hungerford DS. Models of ischemic necrosis of bone. In: Arlet J, Ficat RP, Hungerford DS, editors. Bone circulation. Baltimore: Williams and Wilkins; 1984. p 30-44 
     
    Kawai K, Tamaki A, Hirohata K: Steroid-induced accumulation of lipid in the osteocytes of the rabbit femoral head. A histochemical and electron microscopic study. J Bone Joint Surg Am,1985.67: 755-9, 67755  1985  [PubMed]
     
    Kenzora JE, Steele RE, Yosipovitch ZH, Glimcher MJ: Experimental osteonecrosis of the femoral head in adult rabbits. Clin Orthop,1978.130: 8-46, 1308  1978  [PubMed]
     
    Malizos KN, Quarles LD, Seaber AV, Rizk WS, Urbaniak JR: An experimental canine model of osteonecrosis: characterization of the repair process. J Orthop Res,1993.11: 350-7, 11350  1993  [PubMed]
     
    Brown RE: A surgical approach to the coxofemoral joint of dogs. North Am Vet,1954.34: 420-4, 34420  1954 
     
    Budsberg SC, Verstraete MC, Soutas-Little RW: Force plate analysis of the walking gait in healthy dogs. Am J Vet Res,1987.48: 915-8, 48915  1987  [PubMed]
     
    Budsberg SC, Verstraete MC, Soutas-Little RW, Flo GL, Probst CW: Force plate analyses before and after stabilization of canine stifles for cruciate injury. Am J Vet Res,1988.49: 1522-4, 491522  1988  [PubMed]
     
    Ide T, Hara T, Horii E, An KN, Chao EYS: Prediction of articulating joint pressure distribution under varying loading conditions. Orthop Trans,1991.15: 545-6, 15545  1991 
     
    Palmer JL, Bertone AL, Mansour J: Site specific indentation characteristics of third carpal articular cartilage in exercised versus non-exercised horses. Trans Orthop Res Soc,1993.18: 184, 18184  1993 
     
    Sumner DR, Devlin TC, Winkelman D, Turner TM: The geometry of the adult canine proximal femur. J Orthop Res,1990.8: 671-7, 8671  1990  [PubMed]
     
    Cohen B, Gardner TR, Ateshian GA. The influence of transverse isotropy on cartilage indentation behavior. A study of the humeral head. Read at the Annual Meeting of the Orthopaedic Research Society; 1993 Feb 16; San Francisco, CA 
     
    Emmanual J, Hornbeck C, Bloebaum RD: A polymethyl methacrylate method for large specimens of mineralized bone with implants. Stain Technol,1987.62: 401-10, 62401  1987  [PubMed]
     

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    Anchor for JumpAnchor for Jump
    +Fig. 1:Diagram illustrating the trapdoor technique. The trapdoor is made in the femoral head cartilage (a), after which a 2-cm-diameter area of the femoral head bone is excavated (b). In Group I, the defect was left unfilled (c) and the trapdoor cartilage segment was replaced (d). In Group III, this cartilage segment was collapsed. In Groups II, IV, and V, iliac crest bone graft was harvested (e) and utilized as cancellous bone chips and cortical bone struts (f, g, and h), and the trapdoor was replaced (i). In Group V, osteogenic protein-1 was added.
    Anchor for JumpAnchor for Jump
    +Fig. 2-A:Each femur was secured within a fixation jig fixed to the mechanical testing machine. The fixation jig allowed two degrees of translational adjustment and three degrees of rotational adjustment at the base of the embedded femur. Each test site was aligned with the indentor fixed to the loading actuator.
    Anchor for JumpAnchor for Jump
    +Fig. 2-B:A standard technique was developed to identify the test positions. A circle was mapped onto the femoral head with one point tangential to the plane (plane A) through the posterior edge of the greater trochanter (arrow) and the fovea and a second point tangential to the plane (plane B) initiating at the fovea and perpendicular to the first plane. The test sites were distributed around the perimeter of the circle. For Groups I and II, the posterior-medial (2), anterior-medial (3), anterior-lateral (5), posterior-lateral (6), and center (7) test sites were used. For Groups III, IV, and V, the center site was used again, along with a posterior site (1) and an anterior site (4).
    Anchor for JumpAnchor for Jump
    +Fig. 3-A:Figs. 3-A through 3-E Representative radiographs from the five study groups. Fig. 3-A Group I demonstrated no healing.
    Anchor for JumpAnchor for Jump
    +Fig. 3-B:Group II demonstrated excellent healing.
    Anchor for JumpAnchor for Jump
    +Fig. 3-C:Minimal radiographic healing with collapse of the femoral head was seen in Group III.
    Anchor for JumpAnchor for Jump
    +Fig. 3-D:Moderate radiographic healing without collapse of the femoral head was seen in Group IV.
    Anchor for JumpAnchor for Jump
    +Fig. 3-E:Excellent radiographic healing without collapse of the femoral head was seen in Group V.
    Anchor for JumpAnchor for Jump
    +Fig. 4-A:A histological section through the cartilage of the collapse model (Group III) illustrates the irregularity of the cartilage surface (Goldner trichrome, 25).
    Anchor for JumpAnchor for Jump
    +Fig. 4-B:A histological section of a femoral head treated with bone-grafting (Group IV). There was persistence of the defect where the trapdoor had been made. Fibrous tissue is evident between the ends of the cartilage, where bone has grown underneath the cartilage (Goldner trichrome, 25).
    Anchor for JumpAnchor for JumpTABLE I:  Normalized Stiffness Values*
    *The values are given as the mean and the standard deviation.
    Test SiteGroup IGroup IIGroup IIIGroup IVGroup V
    Anterior-lateral 0.8±0.31.2±0.6
    Anterior-medial0.9±0.31.1±0.6
    Center1.1±0.41.3±0.50.9±0.31.0±0.20.9±0.2
    Posterior-lateral0.9±0.31.0±0.4
    Posterior-medial0.8±0.21.0±0.3
    Anterior1.0±0.20.9±0.21.0±0.4
    Posterior0.9±0.21.1±0.20.8±0.4
    Cornell CN, Salvati EA, Pellicci PM: Long-term follow-up of total hip replacement in patients with osteonecrosis. Orthop Clin North Am,1985.16: 757-69, 16757  1985  [PubMed]
     
    Dorr LD, Takei GK, Conaty JP: Total hip arthroplasties in patients less than forty-five years old. J Bone Joint Surg Am,1983.65: 474-9, 65474  1983  [PubMed]
     
    Katz RL, Bourne RB, Rorabeck CH, McGee H: Total hip arthroplasty in patients with avascular necrosis of the hip. Follow-up observations on cementless and cemented operations. Clin Orthop,1992.281: 145-51, 281145  1992  [PubMed]
     
    Mont MA, Hungerford DS: Current concepts review. Non-traumatic avascular necrosis of the femoral head. J Bone Joint Surg Am,1995.77: 459-74, 77459  1995  [PubMed]
     
    Saito S, Saito M, Nishina T, Ohzono K, Ono K: Long-term results of total hip arthroplasty for osteonecrosis of the femoral head. A comparison with osteoarthritis. Clin Orthop,1989.244: 198-207, 244198  1989  [PubMed]
     
    Stauffer RN: Ten-year follow-up study of total hip replacement. J Bone Joint Surg Am,1982.64: 983-90, 64983  1982  [PubMed]
     
    Buckley PD, Gearen PF, Petty RW: Structural bone-grafting for early atraumatic avascular necrosis of the femoral head. J Bone Joint Surg Am,1991.73: 1357-64, 731357  1991  [PubMed]
     
    Ganz R, Büchler U. Overview of attempts to revitalize the dead head in aseptic necrosis of the femoral head—osteotomy and revascularization. In: Hungerford DS, editor. The hip: proceedings of the eleventh open scientific meeting of the Hip Society. St. Louis: CV Mosby; 1983. p 296-305 
     
    Itoman M, Yamamoto M: Pathogenesis and treatment of idiopathic aseptic necrosis of the femoral head. Clin Immunol,1989.21: 713-25, 21713  1989 
     
    Mankin HJ: Nontraumatic necrosis of bone (osteonecrosis). N Engl J Med,1992.326: 1473-9, 3261473  1992  [PubMed]
     
    Meyers MH, Jones RE, Bucholz RW, Wenger DR: Fresh autogenous grafts and osteochondral allografts for the treatment of segmental collapse in osteonecrosis of the hip. Clin Orthop,1983.174: 107-12, 174107  1983  [PubMed]
     
    Mont MA, Einhorn TA, Sponseller PD, Hungerford DS: The trapdoor procedure using autogenous cortical and cancellous bone grafts for osteonecrosis of the femoral head. J Bone Joint Surg Br,1998.80: 56-62, 8056  1998  [PubMed]
     
    Mont MA, Jones LC, Einhorn TA, Hungerford DS, Reddi AH: Osteonecrosis of the femoral head. Potential treatment with growth and differentiation factors. Clin Orthop,1998.355(Suppl): 314-35, 355(Suppl)314  1998 
     
    Rosenwasser MP, Garino JP, Kiernan HA, Michelsen CB: Long term followup of thorough debridement and cancellous bone grafting of the femoral head for avascular necrosis. Clin Orthop,1994.306: 17-27, 30617  1994  [PubMed]
     
    Smith KR, Bonfiglio M, Montgomery WJ: Non-traumatic necrosis of the femoral head treated with tibial bone-grafting. A follow-up note. J Bone Joint Surg Am,1980.62: 845-7, 62845  1980  [PubMed]
     
    Steinberg ME, Brighton CT, Corces A, Hayken GD, Steinberg DR, Strafford B, Tooze SE, Fallon M: Osteonecrosis of the femoral head. Results of core decompression and grafting with and without electrical stimulation. Clin Orthop,1989.249: 199-208, 249199  1989  [PubMed]
     
    Sugioka Y, Hotokebuchi T, Tsutsui H: Transtrochanteric anterior rotational osteotomy for idiopathic and steroid-induced necrosis of the femoral head. Indications and long-term results. Clin Orthop,1992.277: 111-20, 277111  1992  [PubMed]
     
    Urbaniak JR, Coogan PG, Gunneson EB, Nunley JA: Treatment of osteonecrosis of the femoral head with free vascularized fibular grafting. A long-term follow-up study of one hundred and three hips. J Bone Joint Surg Am,1995.77: 681-94, 77681  1995  [PubMed]
     
    Yamamoto M, Itoman M, Sagamoto N, Morita M: Strut bone graft for aseptic necrosis of the femoral head: theory and surgical technique. Orthop Surg,1983.34: 902-8, Japanese34902  1983 
     
    Wang EA, Rosen V, D’Alessandro JS, Bauduy M, Cordes P, Harada T, Israel DI, Hewick RM, Kerns KM, LaPan P, Luxenberg DP, McQuaid D, Moutsatsos IK, Nove J, Wozney JM: Recombinant human bone morphogenetic protein induces bone formation. Proc Natl Acad Sci USA,1990.87: 2220-4, 872220  1990  [PubMed]
     
    Ficat RP, Arlet J. Functional investigation of bone under normal conditions. In: Hungerford DS, editor. Ischemia and necrosis of bone. Baltimore: Williams and Wilkins; 1980. p 29-52 
     
    Cook SD, Baffes GC, Wolfe MW, Sampath TK, Rueger DC: Recombinant human bone morphogenetic protein-7 induces healing in a canine long-bone segmental defect model. Clin Orthop,1994.301: 302-12, 301302  1994  [PubMed]
     
    Cook SD, Dalton JE, Tan EH, Whitecloud TS 3rd, Rueger DC: In vivo evaluation of recombinant human osteogenic protein (rhOP-1) implants as bone graft substitute for spinal fusions. Spine,1994.19: 1655-63, 191655  1994  [PubMed]
     
    Cook SD, Salkeld SL, Brinker MR, Wolfe MW, Rueger DC: Use of an osteoinductive biomaterial (rhOP-1) in healing large segmental bone defects. J Orthop Trauma,1998.12: 407-12, 12407  1998  [PubMed]
     
    Gerhart TN, Kirker-Head CA, Kriz MJ, Holtrop ME, Hennig GE, Hipp J, Schelling SH, Wang E: Healing segmental femoral defects in sheep using recombinant human bone morphogenetic protein. Clin Orthop,1993.293: 317-26, 293317  1993  [PubMed]
     
    Johnson EE, Urist MR, Finerman GA: Resistant nonunions and partial or complete segmental defects of long bones. Treatment with implants of a composite of human bone morphogenetic protein (BMP) and autolyzed, antigen-extracted, allogeneic (AAA) bone. Clin Orthop,1992.277: 229-37, 277229  1992  [PubMed]
     
    Khouri RK, Koudsi B, Reddi H: Tissue transformation into bone in vivo. A potential practical application. JAMA,1991.266: 1953-5, 2661953  1991  [PubMed]
     
    Lindholm TC, Lindholm TS, Marttinen A, Urist MR: Bovine bone morphogenetic protein (bBMP/NCP)-induced repair of skull trephine defects in pigs. Clin Orthop,1994.301: 263-70, 301263  1994  [PubMed]
     
    Mazières B: Bone morphogenetic protein and bone necrosis: a perspective. ARCO News,1994.6: 3-5, 63  1994 
     
    Mundy GR: Regulation of bone formation by bone morphogenetic proteins and other growth factors. Clin Orthop,1996.323: 24-8, 32324  1996 
     
    Reddi AH: Symbiosis of biotechnology and biomaterials: applications in tissue engineering of bone and cartilage. J Cell Biochem,1994.56: 192-5, 56192  1994  [PubMed]
     
    Riley EH, Lane JM, Urist MR, Lyons KM, Lieberman JR: Bone morphogenetic protein-2: biology and applications. Clin Orthop,1996.324: 39-46, 32439  1996  [PubMed]
     
    Scully SP, Rizk WS, Seaber AV, Urbaniak JR. Augmentation of subchondral bone formation in AVN with rhBMP-2. Poster presentation at the Annual Meeting of the Orthopaedic Research Society. 1995 Feb 13-16. Orlando, FL 
     
    Toriumi DM, Kotler HS, Luxenberg DP, Holtrop ME, Wang EA: Mandibular reconstruction with a recombinant bone-inducing-factor. Functional, histologic, and biomechanical evaluation. Arch Otolaryngol Head Neck Surg,1991.117: 1101-12, 1171101  1991  [PubMed]
     
    Trippel SB, Coutts RD, Einhorn TA, Mundy GR, Rosenfeld RG: Growth factors as therapeutic agents. J Bone Joint Surg Am,1996.78: 1272-86, 781272  1996 
     
    Urist MR: Bone: formation by autoinduction. Science,1965.150: 893-9, 150893  1965  [PubMed]
     
    Urist MR, Dawson E: Intertransverse process fusion with the aid of chemosterilized autolyzed antigen-extracted allogeneic (AAA) bone. Clin Orthop,1981.154: 97-113, 15497  1981  [PubMed]
     
    Wozney JM: Bone morphogenetic proteins. Prog Growth Factor Res,1989.1: 267-80, 1267  1989  [PubMed]
     
    Yasko AW, Lane JM, Fellinger EJ, Rosen V, Wozney JM, Wang EA: The healing of segmental bone defects, induced by recombinant human bone morphogenetic protein (rhBMP-2). A radiographic, histological, and biomechanical study in rats. J Bone Joint Surg Am,1992.74: 659-70, 74659  1992  [PubMed]
     
    Adams RJ. Experimental surgery and physiology: induced animal models of human disease. In: Sindle MM, Adams RJ, editors. Musculoskeletal system. Baltimore: Williams and Wilkins; 1988. p 41-55 
     
    Gold EW, Fox OD, Weissfeld S, Curtiss PH: Corticosteroid-induced avascular necrosis: an experimental study in rabbits. Clin Orthop,1978.135: 272-80, 135272  1978  [PubMed]
     
    Jones LC, Hungerford DS. Models of ischemic necrosis of bone. In: Arlet J, Ficat RP, Hungerford DS, editors. Bone circulation. Baltimore: Williams and Wilkins; 1984. p 30-44 
     
    Kawai K, Tamaki A, Hirohata K: Steroid-induced accumulation of lipid in the osteocytes of the rabbit femoral head. A histochemical and electron microscopic study. J Bone Joint Surg Am,1985.67: 755-9, 67755  1985  [PubMed]
     
    Kenzora JE, Steele RE, Yosipovitch ZH, Glimcher MJ: Experimental osteonecrosis of the femoral head in adult rabbits. Clin Orthop,1978.130: 8-46, 1308  1978  [PubMed]
     
    Malizos KN, Quarles LD, Seaber AV, Rizk WS, Urbaniak JR: An experimental canine model of osteonecrosis: characterization of the repair process. J Orthop Res,1993.11: 350-7, 11350  1993  [PubMed]
     
    Brown RE: A surgical approach to the coxofemoral joint of dogs. North Am Vet,1954.34: 420-4, 34420  1954 
     
    Budsberg SC, Verstraete MC, Soutas-Little RW: Force plate analysis of the walking gait in healthy dogs. Am J Vet Res,1987.48: 915-8, 48915  1987  [PubMed]
     
    Budsberg SC, Verstraete MC, Soutas-Little RW, Flo GL, Probst CW: Force plate analyses before and after stabilization of canine stifles for cruciate injury. Am J Vet Res,1988.49: 1522-4, 491522  1988  [PubMed]
     
    Ide T, Hara T, Horii E, An KN, Chao EYS: Prediction of articulating joint pressure distribution under varying loading conditions. Orthop Trans,1991.15: 545-6, 15545  1991 
     
    Palmer JL, Bertone AL, Mansour J: Site specific indentation characteristics of third carpal articular cartilage in exercised versus non-exercised horses. Trans Orthop Res Soc,1993.18: 184, 18184  1993 
     
    Sumner DR, Devlin TC, Winkelman D, Turner TM: The geometry of the adult canine proximal femur. J Orthop Res,1990.8: 671-7, 8671  1990  [PubMed]
     
    Cohen B, Gardner TR, Ateshian GA. The influence of transverse isotropy on cartilage indentation behavior. A study of the humeral head. Read at the Annual Meeting of the Orthopaedic Research Society; 1993 Feb 16; San Francisco, CA 
     
    Emmanual J, Hornbeck C, Bloebaum RD: A polymethyl methacrylate method for large specimens of mineralized bone with implants. Stain Technol,1987.62: 401-10, 62401  1987  [PubMed]
     
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